1992 PhD “Nutrient dynamics and nitrogen trace gas flux during ecosystem development in montane rainforest” Stanford University, USA
1982 BSc Forestry, University of Washington, USA.
2002-11 Western Washington University, USA. Adjunct faculty.
2002-06 Earthwatch Institute, USA.Field Director.
2000-02 Western Washington University, USA. Director, Off-Campus Environmental Studies.
1996-00 University of Otago, USA. Post-doctoral Research Fellow.
1993-94 National Council for Air and Stream Improvement, USA. Forest Scientist.
My research interests include:
Interactions between species and how these are modified by environmental conditions
Effects of climate change, invasive species and environmental disturbances on ecosystems
Land-use effects on freshwater systems
I am course co-ordinator for:
BI201 General Ecology
BI206 Quantitative Biology
Dolédec, S., N. Phillips, M. Scarsbrook, R.H. Riley, & Colin R. Townsend (2006).Comparison of structural and functional approaches to determining landuse effects on grassland stream invertebrate communities.Journal of the North American Benthological Society 25:44-60.
Riley, R.H., C.R. Townsend, D. Raffaelli & A. Flecker (2004) Sources and effects of subsidies along the stream-estuary continuum. In Food webs at the landscape level.Editors: G.A. Polis, M.E. Power and G.R. Huxel.University of Chicago Press, Chicago, IL.Pages 241-260.
Riley, R.H. & R.L. Jefferies (2004) Subsidy dynamics and global change. In Food webs at the landscape level.Editors: G.A. Polis, M.E. Power and G.R. Huxel.University of Chicago Press, Chicago, IL.
Riley, R.H., C.R. Townsend, D.K. Niyogi, C.A. Arbuckle & K.A. Peacock (2003) Headwater stream response to grassland agricultural development in New Zealand. New Zealand Journal of Marine and Freshwater Research 37:389-403.
Huryn, A.D., R. Riley, R.G. Young, K. Peacock, C.J. Arbuckle (2002) Natural-abundance stable C and N isotopes indicate weak upstream-downstream linkage of consumer food webs in a river-floodplain system. Archiv für Hydrobiologie 153:177-196.
R.M.Thompson, C.R.Townsend, D.Craw, R.Frew and R.Riley (2001) Further links from rocks to plants: potential for ectomycorrhizal fungi to mobilize nitrogen from minerals.Trends in Ecology and Evolution 16:543.
Hall, M.J., Closs, G.P. & R.H. Riley (2001) Relationships between land-use and stream invertebrate community structure in a South Island, coastal stream catchment.New Zealand Journal of Marine and Freshwater Research 35: 591-603
Huryn, A.D., R.H. Riley, R.G. Young, C.J. Arbuckle, K. Peacock, and G. Lyon (2001) Temporal shift in contribution of terrestrial organic matter to consumer production in a grassland river. Freshwater Biology 46:213-226.
Riley, R.H. & P.M. Vitousek (2000) Hurricane effects on nitrogen trace gas emissions in Hawaiian montane rainforest.BioTropica 32:195-199.
Townsend, C.A. & R.H. Riley (1999) Assessment of river integrity - accounting for perturbation pathways in physical and ecological space.Freshwater Biology 41:393-405
Riley, R.H., C. Mitchell & J. Thorp (1998) UV and stream ecology. In UV radiation and its effects: an update. National Science Strategy Committee for Climate Change Workshop, Christchurch, December 1997. The Royal Society of New Zealand. Miscellaneous Publications 49.1998.
Griffiths, R.P., P.S. Homann, and R. Riley (1998) Denitrification enzyme activity of Douglas-fir and red alder forest soils of the Pacific Northwest. Soil Biology & Biochemistry 30:1147-1157.
Potter, C.S., R.H. Riley & S.A. Klooster (1997) Simulation modeling of nitrogen trace gas emissions along an age gradient of tropical forest soils.Ecological Modeling 97, 179-196.
Riley, R.H., M. Schuft, D.L. Phillips & M.C. Garcia (1997) Resolution and error in measuring land cover change: effects on estimating net carbon release from Mexican terrestrial ecosystems.International Journal of Remote Sensing 18(1):121-137.
Riley, R.H. & M. Schuft(1995) Mexico Case Study In Cairns, M, A., T.P. Kolchugina, D.P. Turner & J.K. Winjum. The Contribution of Forest Land Use to Total National Carbon Flux: Case Studies in the Former Soviet Union, United States, Mexico, and Brazil.US EPA EPA/600/R-95/044.
Riley, R.H. (1996) Process-level regulation of nitrogen trace gas flux in Hawaiian montane rainforest Soil Biology and Biochemistry 28:1251-1260.
Crews, T.E., K. Kitayama, J.H. Fownes, R.H. Riley, D.A. Herbert, D. Mueller-Dombois & P. M. Vitousek (1995) Changes in soil phosphorus fractions and ecosystem dynamics across a long chronosequence in Hawaii. Ecology 76:1407-1424.
Riley, R.H. & P. M. Vitousek (1995) Nutrient dynamics and nitrogen trace gas flux during ecosystem development in montane rainforest.Ecology 76(1):292-304.
Raich, J.W., R.H. Riley & P.M. Vitousek (1994) Use of root ingrowth cores to assess nutrient limitations in forest ecosystems.Canadian Journal of Forest Research.24:2135-2138.
Davidson, E.A., P.A. Matson, P.M. Vitousek, R. Riley, K. Dunkin, G. García-Méndez& J. M. Maass (1993) Processes Regulating Soil Emissions of NO and N2O in a Seasonally Dry Tropical Forest.Ecology 74(1):130-139.
Davidson, E.A., P.M. Vitousek, P.A. Matson, R. Riley, G. Garcia & J.M. Maass (1991)Soil Emissions of Nitric Oxide in a Seasonally Dry Tropical Forest of Mexico. Journal of Geophysical Research 96:15439‑15445
Riley, R.H. (2010) Report to the USFS and Tulalip Tribes on the Segelsen Ridge Huckleberry Enhancement Response Monitoring Report.
Riley, R.H. (2004-06)Report to the USFS and Skagit Fisheries Enhancement Group on stream channel surveys and riparian conditions in the Skagit River Basin
Riley, R.H. & D.W. Cole (1995) Report to the Superintendent of Wastewater Facilities, City of Bremerton, on five years of forest growth following biosolids application. University of Washington